# Species Richness & Evenness (S, Margalef, Menhinick, J = H′/H′max)

Richness S with Margalef/Menhinick indices, Shannon H′, Pielou evenness J, and Berger–Parker dominance from a named or plain species list.

> Canonical page: https://elysiatools.com/en/tools/species-richness-evenness

- **Category:** Health

- **Keywords:** species richness, pielou evenness, margalef index, menhinick index, berger-parker dominance, community structure, biodiversity calculator, ecology calculator, quadrat sampling, species abundance, shannon evenness, field ecology

## Overview

Paste a named species list or plain abundance counts to compute richness S, sample-size-corrected Margalef and Menhinick indices, Shannon H′, Pielou evenness J = H′/H′max, and Berger–Parker dominance of the most abundant taxon.

## Inputs

- **Species counts (one per line, `name: count` or plain numbers)** (textarea): One species per line as `name: count` (separators : ; = , | or tab), or plain numbers separated by new lines, commas, or spaces. Duplicate names are merged by summing.

## When to use

- After quadrat or plot sampling when you need S plus evenness and dominance in one step.
- To compare communities of different sample sizes using Margalef d = (S−1)/ln N and Menhinick d = S/√N.
- When abundances look skewed and you want Pielou J and Berger–Parker to quantify how uneven the assemblage is.

## How it works

- Enter one species per line as name: count (separators : ; = , | or tab) or as plain numbers separated by newlines, commas, or spaces; duplicate names are summed.
- The tool totals individuals N, counts unique species S, and identifies the dominant taxon.
- It reports Margalef d, Menhinick d, Shannon H′, H′max = ln S, Pielou evenness J, and Berger–Parker dominance (proportion of the most abundant species).

## Use cases

- Summarizing tree or herb counts from forest quadrats into richness, evenness, and dominance.
- Classroom or lab exercises that contrast even versus dominated assemblages from the same N.
- Rapid comparison of community structure between sites when only a species-abundance list is available.

## Frequently asked questions

### What input formats are accepted?

Named lines such as Oak: 34 (or with ; = , | or tab) or plain numbers on separate lines or separated by commas or spaces. Duplicate names are merged by summing counts.

### How is Pielou evenness J calculated?

J = H′ / ln S (i.e., H′ / H′max). Values near 1 indicate nearly equal abundances; lower values indicate dominance.

### What do the Margalef and Menhinick indices measure?

Both correct richness for sample size: Margalef d = (S − 1) / ln N and Menhinick d = S / √N. Higher values mean more species relative to the number of individuals counted.

### Can I submit unnamed counts?

Yes. Plain numbers are treated as successive species (Species 1, Species 2, …) so dominance can still be reported.

### What is Berger–Parker dominance?

The proportion of all individuals that belong to the single most abundant species. High values (e.g., 0.8) indicate a strongly dominated community.

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